Just from looking at the still of the video above, as I’m at work and can’t watch it, it also looks like they are using 3 LFO modules (well at least 1 for sure) to modulate the granular module. I’ll check out the video later today, and see if I can sleuth anymore.
Edit: yep precisely what they are doing. Looks like the granular is being used with external audio and is then being filtered and output. The lfos involved are being used to modulate the granular module’s parameters.